A medical doctor, naturalist, and wildlife photographer. Member of the American Society of Mammalogists and the Spanish Society for Conservation and Study of Mammals.
Author of Bovids of the World: Antelopes, Gazelles, Cattle, Goats, Sheep, and Relatives (Princeton), and Canids of the World: Wolves, Wild Dogs, Foxes, Jackals, Coyotes, and Their Relatives.