The integration of Harold and Primrose with Brindabella and her calf was accidental. The former pair broke down a barrier dividing the exhibit.
The barrier wouldn't have been more than a wooden fence or something along the lines of that. One side of the original ponds can be seen on the Safari tour; to the right of the drivethrough pond.
We can assume the previous groupings (at the time of 1999) were; Beatrice, Brindabella and Genevieve and Primrose, with Harry off display. I'd assume Makoko was probably imported, due to space opening up following the passing of Genevieve, and so would've been grouped with Primrose. Harry may have then been moved in with Beatrice and Brindabella to breed. Beatrice died at a young age, so it's likely she may have died during childbirth.