Interesting that the 'natural only- fail ' actually means 'no mating at all'(!) This Adelaide pair represent another depressing catalogue of failure, so similar to some other zoos. You are correct, a few pairs 'click',are obviously compatabile, mate normally and breed repeatedly. Or else- as in the majority of pairs, there is no breeding at all. Still nowhere seems to take into account the 'lek' behaviour in the wild in this species, allowing both stimulation/rivalry between males and the female to choose from several males which one she will actually mate with. Such a choice is completely denied in captivity, at least outside China. Not surprising the failure rate is so high. Why are they all so slow to recognise this and try to adjust breeding attemps to better mirror the wild behaviour? Presumably because the zoos are all tied by China only allowing pairs at a time to be loaned, and never more than that. But while that continues, so will the the problem of achieving a more successful breeding rate.
In most cases were natural only mating was attempted, the pandas didn’t even mate, so there was no chance of pregnancy. The majority of the births in the US were via AI, which makes the cubs born via natural mating at the National Zoo to their original pair all the more remarkable.
I’ve seen comments from Adelaide Zoo that detail Funi’s willingness to mate and Wang Wang’s reluctance. It appears his performance is the issue here and could indeed be improved with a stimulant male. Since the males would never meet face to face in captivity, I see no reason why the scent of another male (acquired from another zoo) couldn’t be used to achieve the same result.
As with the great apes back in the early-mid 20th century, I suspect the general public are still under the anthropomorphic impression pandas live in couples.