The 'article' I quoted: -
VERVAECKE (H.), STEVENS (J.) & VAN ELSACKER (L.), 2004 : Pan continuity : bonobo-chimpanzee hybrids. Folia Primatologica 75(1): 59
is in fact just a short abstract of a presentation given at the Spring 2003 meeting of the Primate Society of Great Britain.
The abstract reads:
From historical accounts it appears that natural populations of bonobos and chimpanzees have been allopatric since their phylogenetic separation. There are no accounts of hybridisation under natural conditions. There is, however, evidence for interbreeding between bonobos and chimpanzees in captivity. We briefly (6.5 h) observed and filmed a small group of four bonobo-chimpanzee hybrids (2 males, aged 10 and 9, and 2 females aged 10 and 8).
There was clear individual variation in degree of expression of typical chimpanzee or bonobo features. In each individual, there were anatomical features reminiscent of both of the parental species: the specific bonobo-like pink coloration of the lips, the hairstyle and slender body build, the chimpanzee-like dorsal position of the female genitals, more sturdy body build and prognathism. The behaviours could be categorized less exclusively. We observed mostly female oriented interactions, including grooming, play, approaches and intersexual ventro-dorsal sex. There was one female coalition against the males. The individuals lip-smacked while grooming and performed swaggering displays. The hoots were like chimpanzee pant-hoots with an introduction, build-up and climax in a higher pitched bonobo tone and more e than o sounds. Studies on Pan in the seventies and eighties emphasised discontinuity between the two species, contrasting the male-dominated, aggressive nature of the
chimpanzee to the female-oriented, peaceful nature of the bonobo. In the nineties and in present studies, the continuity among Pan is increasingly being documented. The existence of hybrids challenges our tendency for binary thinking and points at the conceptual relativity of the species gap.