Let's revisit Darwin's finches for a moment. Island adapted species are often extreme ecological specialists. There are more than one species of rail present and any number of Ardeidae-each bird has its own special adaptations -foraging on organisms perfectly dispatched and ingested with the least expenditure of energy -consequent of bill shapes and foraging behaviors, hours of activity, hours of respite- -islands can be hotspots of endemism because of the tendency for its inhabitants to become extreme ecological specialists. They're not delicate - quite the opposite- they're resilient-
I'm not quite sure of the relevance of this passage. We aren't talking about insular life in general, we are talking about one particular species on one particular island group: banded rails are in no way comparable to Galapagos finches. On the Cocos (Keeling) Islands there are exactly two species of rails established as breeders, the banded rail and the white-breasted waterhen, which have exclusive life-styles. The only established Ardeidae are nankeen night herons and reef herons, also both exclusive to the life-styles of both of the rails. All of these are widespread species in the Pacific area, of which only one (the banded rail) has an endemic subspecies on the Cocos (Keeling) Islands.
Specialist insular species are not resilient....if that were the case there would not be so many extinct specialist insular species. When generalist species are introduced by man, the specialist species are forced out. In other words specialist species do very well when their ecosystem is intact, but fare badly when their ecosystem is corrupted.
As it happens, banded rails are not specialists in any way, they are generalists. The junglefowl are not the only threat to their survival on the islands, but they are one of them (i.e. the rails are not endangered solely because the junglefowl were introduced, but junglefowl are one of the combatable factors of the rails' endangerment).
mihos said:
We're quickly adopting postions for or against the survival of Cocos Junglefowl- when we might be better served discussing endemism and island biogeography.
that is what the thread is about. The banded rail subspecies of the Cocos (Keeling) Islands is endemic to those islands, while the junglefowl (of both species) are introduced and compete for the same resources. The junglefowl are one of the manageable threats to the endemic bird.
mihos said:
Islands attract pioneers- island hopping species are sometimes introduced to islands in efforts to conserve them. The green junglefowl is an island hopping species adapted to survive on oceanic islands. It didn't reach the Cocos by flying. That can't be debated.
again, I'm not sure what you're saying here: green junglefowl live on islands in the Indonesian archipelago, and therefore introducing them to oceanic islands where they don't belong is acceptable for that reason, and they should remain there even if they have been shown to be a threat to the endemic life-forms of that island?
mihos said:
Wild populations of Red Junglefowl- introduced by early seafarers- on islands across the Pacific and Oceania are/were cohabitants with terrestrial rails. Many of these rails are endemic- and unfortunately many species are extinct due to the introduction of rats and pigs. But it's unlikely wild Junglefowl are serious competitors- Their life histories don't overlap closely. Rails are generally active at hours the junglefowl is not. The rails specialize on certain invertebrates, mollusks and shoots of vegetation, green junglefowl are not adapted for. And rails enjoy eating the eggs and chicks of other terrestrial birds. While rails are omnivores- compared with junglefowl-are obligatory predators. The organisms preferred by rails are in an arms race with their prey. this arms race takes place in the densest vegetation- often marshy vegetation- and during hours when aquatic lifeforms are most visible /active. The junglefowl is never going to compete with the rail in its preferred- in its optimal habitat and neither is the rail competition for the junglefowl.
again, you are speaking in extremely general terms when the topic at hand is solely about junglefowl/banded rails on the Cocos (Keeling) Islands. If you were going to talk generally you should have also mentioned that diseases introduced with domestic fowl and other gamebirds are generally accepted as being the cause or probable cause of extinction or endangerment for a number of island birds, including the New Zealand quail (extinct) and weka (endangered). But introduced disease is apparently not a factor on the Cocos (Keeling) Islands, and thus is of no more relevance than what the factors are at play on other island groups. Your last comments are a nonsense with regards to the life-style of banded rails: they are cathemeral generalists, not (if I am reading you right) marsh-dwelling nocturnal specialists. The fact is they feed on the same food items that junglefowl take, and during the same periods (as well as at night).
mihos said:
Rails have such distinctive morphological features-that make them perfectly adapted for life on remote islands- together with other species- a diversity of species- Diversification is the key to life-
The natural history of both species is worthy of serious investigation by objective researchers and that island biogeography - that Alfred Wallace are revisited- and utilised as more important reference for conservationists.
and if I'm reading this right, you are basically saying that introducing foreign species to insular ecosystems is a good thing? Because it increases the biodiversity? I think island biogeographers might have different ideas about what the outcome of that might be.....
Just some additional information for specifics: the banded rail subspecies found in these islands is now probably confined solely to North Keeling Island which is only 1.2 km² in size. It has been eliminated from all the other islands in the group through a combination of habitat destruction (land clearance) and introduced predators (rats and cats in particular). Competition with junglefowl has also been proven to be a factor. Yellow crazy ants are another major threat now (as is also the case on Christmas Island for the endemic land crabs: nasty things those ants!). Clearing the other islands of introduced species (e.g. rats, cats, junglefowl) will enable rails to be reintroduced there for an increase in the overall population.