To add a new (related) discussion topic, how can densely urban zoos expand? Not just London Zoo, but any zoo in a large city with little open space?
Back to this sub-topic.
It is possible to put grass lawns and trees inside animal exhibits as 'green islands'. Ponds can become animal pools or water moats. One can plan service area so that access roads and corridors are minimized. Avoid nonsense public plazas. Ornamental plantings, sculptures, theming - this all can go inside animal exhibits, eventually protected by hotwire or elevated. Hide night stalls and service buildings under the paddocks. Perhaps you shouldn't let a herd of buffalo on the roof, but small and medium-sized animals, birds, carnivores, smaller hoofstock etc. can run on the green roof of any building without major structural adjustments.
A leader is probably small zoo in Bremenhaven, Germany, which was redesigned. In Bremenhaven, the combined size of animal exhibits is bigger than the total zoo area. Eg. the aquarium and restrooms are under the rocks of polar bears. Hill-shaped nature of many exhibits betrays that service buildings and night stalls are below them. Zoo classroom is above restaurant, and visitor paths are often two-level - upper overlooks the exhibit, and lower underwater view of it.
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