Where do the Deer & Antelope Play?: A Look at America's Ungulate Populations

I believe that some of the private collections within the US are quite large numbers!

Yes there are a lot of populations outside of the AZA which are managed by private keepers. In fact, there are a good number of species managed privately which aren't in public zoos at all anymore. Nilgiri Tahr, Topi, and Grey Rhebok are all examples, though I can't say I know how well those populations are doing.

Excelent thread !
I am curious about the institutions, hold Mexican Red Brocket. Gladys Porter , Huston and ?

Thanks! :)

Gladys Porter Zoo in Texas and Bergen County Zoo in New Jersey are currently the only AZA holders of brocket deer. The other two holders are non-AZA but I do not know who they are or if they are public.

Great work Thylo, really interesting. Two things leapt out at me from the list though -

That's a lot of Anoas across not many zoos. Are some of those also held privately?


That's one hell of a drop in numbers.

Also, no Southern Black Rhino figures? Thought they were a tad more common over there than in Europe?

Thank you! Good a good deal of time but I'm happy people are interested.

In the 2014 report, those 0.0.70 anoa all belonged to one of three private holders. I'd imagine a good deal of the current population still remains in private hands but is managing them in cooperation with the TAG.

I meant to make a note on the Gerenuk drop so thanks for reminded me! While I do not know what caused the dropped, it was a lot more sudden than my post suggests sadly. In 2017, there were 37.47 Gerenuk in the population. However, in the 2018 report there were only 22.29 left. Over the course of one year the population lost 15.18 animals... The good news is that the population has at least slightly increased between 2018 and 2019, even if only by 1.3 animals. Despite the sudden drop in numbers, the TAG still lists the species as a Yellow SSP and considers the population to be stable.

No unfortunately. I don't think the South-Central Black Rhinoceros program was ever an AZA program, but rather one managed by Fort Worth which other zoos joined on their own accord. I think the program has been deemed unsustainable now, unfortunately, and the taxa is listed for phasing out. I know Fort Worth, Disney, White Oak, and Denver(?) still hold the subspecies, though I don't know if there are any others or what their numbers are.

Why is the endangered Calamian deer being phased out?

All the females died. LA kept breeding them right to the end but I believe all new fawns born and eventually much of the herd (possibly all of the herd by now?) succumbed to chronic wasting disease.

@ThylacineAlive I want to thank you for posting this thread, I’ve been getting more and more into ungulates in the past year due to being able to visit zoos here in California. It’s awesome to seem massive mixed species exhibit in the likes of the San Diego Zoo Safari Park mingling like it’s just a group of people out at brunch :D I hope to see the Calamian Deer at the LA Zoo next week, they almost always are ignored by average zoo goers but find it sad as well as @Gavialis mentioned. I’ll want to ask the hoofstock keepers if I manage to see one about their situation.

You are very welcome!

If you do manage to find Calamian Deer still at LA, please let me know how many you see, or try to find out how many are left if you do speak to a keeper. Besides LA, the only other holder will be Phoenix with either one or two males.

This is an extremely interesting thread; thanks for starting it.

I note that royal antelope doesn't feature on your list. I saw one at Lowry Park in 2009; when was this species last kept in the USA?

(The royal antelope I saw at Lowry Park was the first I'd seen since the species was kept at London Zoo when I was a child.)

My pleasure :)

I, too, saw the pair of Royal Antelope at Lowry Park in 2013 (the only time I've seen the species). If they're still around, it's down to 1.0 kept at San Diego Zoo. I'm not entirely positive this individual is still alive, however...

Given that the number of zoos holding mountain zebra doubled in 5 years and the population increased by over a third, it seems like that has indeed been happening. The drop in plains zebra has been more precipitous than can probably be explained by that, though.

This does happen and it's indeed a shame, but it's worth noting that the situation is more nuanced than that for ungulates, and for deer in particular. Biosecurity protocols related to ungulates and livestock in the US have led to very restrictive import laws. For deer, the presence of chronic wasting disease in our wild populations has led to a complete ban on import of deer from virtually every other country as well as severe restrictions on moving them across state borders. Frankly, the fact that a few exotic deer programs are still hanging on is already remarkable.

Additionally, Plains Zebra are incredibly common in the private trade and as an exotic pet. I could buy one online right now if I wanted to. As such, they are a very common animals in non-AZA zoos, safari parks, and ranches. I even visited a ski resort once that kept a Plains Zebra with a camel in their horse barn. Unfortunately, I'm not too sure what the prospects of pure Plains Zebra subspecies are. I've discovered recently that a great deal of the Grant's in US zoos (including AZA) have at least a little hybridization with Burchell's in their ancestry. As for Burchell's or any others, who knows.

It's quite sad how badly captive deer populations have suffered due to CWD restrictions, though I agree it is remarkable how many programs are still surviving. The fact that the Barasingha, Pere David's Deer, Southern Pudu, and even muntjac programs have grown is excellent news overall. I know muntjac in particular are often one of the species singled out by CWD legislation which is probably a bit surprising given how common they are both in zoos and even in the private trade.

Incidentally, CWD restrictions are something I was just looking into yesterday as well. Simply put, there are only 2 US states that have no written laws specifically referring to CWD. These are Nevada and Washington*, who, in general, banned the keeping of captive deer for seemingly unrelated reasons before CWD restrictions started being implemented in 2002. Additionally, there are 32 states that have some form of regulation that restricts the importation, transfer, and/or keeping of captive cervids to varying degrees of severity. The most common regulation I found was states requiring a 5 year quarantine period of all animals planned to be moved across state lines as well as for all other animals kept in the herd. The remaining 16 states in the US have implemented a ban on the importation (and sometimes even the keeping) of captive cervids to some degree. Some states only ban certain species (native species and all Cervus and Muntiacus species tend to be the most universally banned, though sometimes muntjacs are permitted), though most states ban all cervids from moving into their state lines. Many states also require testing and permitting to even move deer within the state.

As you can imagine, all of these restrictions make it extremely difficult to manage deer programs, especially when these restrictions also pertains to simply transporting deer through states which inhibits zoos' abilities to transport animals between them for breeding. For example, I know the newest legislation passed in Louisiana has meant that the Audubon can no longer ship their Barasingha out of state, meaning the portion of the population kept between the zoo and breeding center are trapped there until further notice.

It's no surprise really that most zoos have simply given up on keeping exotic deer as a result of all this, as sad as that is. I know California and New York have particularly lax (and in my opinion very sound) regulations, both allowing animals planned for import to be tested immediately prior to transport as well as upon arrival in a quarantined setting, so this is why one will find the two major hubs of Asian deer in America to be the Bronx Zoo and the San Diego Zoo Safari Park. These two zoos almost single-handedly manage multiple Asian hoofstock populations between them, including the non-AZA North Indian Muntjac program. I know Bronx alone has roughly half the Barasingha in the United States.

*Why Northwest Trek is permitted to keep Roosevelt Elk, a species high on the CWD threat list is unknown to me. Washington's ban on captive cervids was passed in 1993.

Not sure of how much that phenomenon has contributed to declines in ungulate taxa, although my suspicion is that it has had a sizable impact. Popular flagship species (like giraffes, zebras, and rhinos) and African ungulates who do well in mixed-species savanna habitats seem to have fared pretty well in recent years, while most Asian hoofstock, deer, caprids, and some wild pigs have suffered. However, there are also zoos with designated ungulate areas that have not reduced or replaced those exhibit areas, but have noticeably reduced their number of ungulate taxa - so I suspect other factors like development of formal population management and increase in the use of flex space have contributed to these declines as well.

In addition to these points, many zoos building new African savannas often either do not have the space to accommodate large mixed plains with multi-species herds and therefore prioritize the most crowd-pleasing species, or simply have no interest in a wide diversity of African antelope/gazelle. Simply put, I'd imagine most zoos with more limited holding capacity will have an "any antelope will do" mentality, if they even go for any antelope species at all. While there are still some zoos with designated hoofstock exhibits, as with the small mammal house and carnivore house these exhibits are becoming increasingly endangered in their own right. This greatly reduces the space for ungulate diversity within a collection, as a zoogeographic exhibit will often include many non-ungulate species. It does seem that African hoofstock has still faired better than Asian by and large, though. I'd imagine the interest in large mixed African savannas is much greater than Asian ones. It's possible that African antelopes are seen as more charismatic than the handful of Asian ones present (apart from Arabian Oryx I suppose, but that suffers from having to compete with three other oryx programs). Seeing as deer and caprines are the most populous Asian hoofstock in captivity while also being the least popular with the average zoogoer, it's no surprise to me that large African antelopes have garnered more interest regardless of legislation.

I think the biggest contributing factor will be the inability to import additional stock, however. Much like deer, the USDA has place strict restrictions on the importation of many ungulate groups from out of country. Deer can only be imported internationally from Canada, subject to the CWD restrictions present in both the originating province and the receiving state. As far as other ungulates go, it is nearly impossible to import any ruminant ungulates from Europe, Asia, or Africa. For those unfamiliar with the term, ruminant ungulates are those which have specialized stomachs which ferment their food prior to digestion. Pronghorn, chevrotains, giraffes, musk-deer, true deer, and all of Bovidae (cattle, caprines, antelope, etc.) are ruminants, and therefore subject to importation bans. Imports from Canada are possible, as are (oddly) imports from specifically Australia and New Zealand. As we all know, however, these are not ungulate-rich countries and therefore these exceptions are of little help to US programs. These laws exist to protect the American livestock population of potential diseases transmitted by foreign ruminants. The unfortunate result of these laws is that it is nearly impossible for US zoos to import additional, and often badly needed, bloodlines to their populations. As sad truth for most zoo managed populations is that they often times have a very small founder base. American populations for species like the Slender-Horned Gazelle and North Sulawesi Babirusa, for instance, all descend from three founding animals. Some animals don't suffer from inbreeding depression too much, if at all, but in ungulates the result can often be increased infant mortality and reduced lifespans. Fertility may also be reduced, though I'm not sure if this is as large an issue for US programs as the first two are. While some species have managed to thrive relatively well on low genetic diversity (the aforementioned gazelle and babirusa for example), many others have faced difficult breeding and management challenges as a result. Often times, lack of genetic diversity and its effect on the captive population is as much a factor in determining which programs to phase-out as interest in a species is. The ungulate TAGs work very hard to overcome these challenges and import further genetics whenever possible, but unfortunately the inability to simply bring in more animals handcuffs many programs.

Thanks for all this effort! It's encouraging to see some of the increases here, like addax and pygmy hippopotamus.

An interesting thing to note: many of the species recently being pushed as "priority" by the AZA's Ungulate TAG are from these geographic areas.

Ungulate Profiles — AZA Ungulates

I don't know how much sway the TAG's recommendation to hold species has, but it's nice to see these little profiles espousing the merits of some of these animals + in-situ conversation opportunities :)

You are welcome :)

Thank you for linking those profiles, as it actually gives me figures for four more species:

Guanaco
2017: 34.36.13 in 26 AZA institutions

Domestic Bactrian Camel
2018: 46.55 in 34 AZA institutions

Ugandan Kob
2019: 12.24.2 in 2 institutions

American Bison (pure+impure)
2019: 93.186.10 in 49 AZA institutions

~Thylo
 
I know from second hand experience Bongo, Scimitar, and the various Goat species are breeding like rabbits in private collections. I've seen recently that generic Wildebeest numbers (I assume Blue based on phenotype) are increasing in the South as well. But that was from almost none five years ago.

Bongo and Scimitar Horned Oryx are species that I constantly see for sale. They're not terribly expensive either.

That's a lot of Anoas across not many zoos. Are some of those also held privately?

Yes. There is a privately owned zoo in Wisconsin that has Anoa and currently plans on acquiring more.





I wasn't familiar that Guanacos were so common in the United States. Which facilities have them?
 
Bongo and Scimitar Horned Oryx are species that I constantly see for sale. They're not terribly expensive either.



Yes. There is a privately owned zoo in Wisconsin that has Anoa and currently plans on acquiring more.





I wasn't familiar that Guanacos were so common in the United States. Which facilities have them?

Hunting a Bongo though.still costs as much as a Range Rover. Whereas Scimitar basically are the same price as a Whitetail.

But that was why I deleted my last post. Didn't want to derail a thread about the AZA with private facility talk.
 
You didn't consult me for this :p I'm doing ungulates as soon as I finish carnivores, which should be soon, ideally within a week. It'll take me a while to compile the lists and then proof them as best as possible, but if there's any particular species people would like me to do first, just ask.
 
I wasn't familiar that Guanacos were so common in the United States. Which facilities have them?
This isn’t a complete list, but zoos that have Guanacos include (can’t 100% confirm that some of these places still hold them): San Diego, Detroit, Rosamond Gifford, San Francisco, Louisville, Santa Ana, Capital of Texas and Bergen County.

I actually thought there were more Guanacos in the US than 80 or so individuals, although that might be because of how similar to Llamas:D (although I do like them quite a bit more)
 
I wasn't familiar that Guanacos were so common in the United States. Which facilities have them?

I haven't proofed this list because I can't really tell the difference between guanaco and llama, but:
Abilene, Alabama Gulf Coast, Alameda Park, Audubon, Bergen County, Capital of Texas, Detroit, El Paso, Fairview Zoo (IA), Franklin Drive Thru Safari (TX), Gone Wild Safari (LA), Idaho Falls, Ivie Acres Farm (UT), Louisville, Miller Park, Rosamond Gifford, San Francisco, Santa Ana, Santa Fe College Teaching Zoo, Smoky Mountain Deer Farm (TN), Tennessee Safari Park, Wildlife Safari (OR), Zoo Miami.

All have 1-5 except for TN Safari Park, which has around 30.

Regarding Anoa, they are starting to turn up in roadside zoos that purchase from the pet/ranch industries, so there must be at least one private holder, if not more.
 
I think that the effects of Ernest Hemingway having been lost to us 60 years ago, include a rising disinterest in “Game Species” and an lessening appreciation for their diversity. *half-kidding, but I do believe it to be a factor. Fortunately for now, drive through safari parks pick up some of the slack.
 
Regarding Anoa, they are starting to turn up in roadside zoos that purchase from the pet/ranch industries, so there must be at least one private holder, if not more.

Timbavati Wildlife Park in Wisconsin Dells, Wisconsin has some. I'm not aware of how many they have, as I haven't been there in years, but I'm acquaintances with the child of the owners, and he said they planned on getting more from another facility.
 
That would be great to add the non-AZA numbers as there are large collections of many endangered species in them!

I would love to, but that information will likely be harder to attain. I do plan on starting species lists soon, though, and hopefully members can further add to them. I'll probably do separate posts by "type" of ungulate so as not to have one massive post.

~Thylo
 
Giraffes
Masai Giraffe (10 holders)
Brevard Zoo
Cincinnati Zoo & Botanical Garden
Franklin Park Zoo
Houston Zoo
Los Angeles Zoo & Botanical Gardens
Nashville Zoo
Roger Williams Park Zoo
San Diego Zoo
San Diego Zoo Safari Park
Santa Barbara Zoo

Rothschild's/Nubian Giraffe (suspected) (2 holders)
B. Bryan Preserve
San Diego Zoo Safari Park

Giraffe (sensu lato) (19 holders)
Audubon Species Survival Center
Audubon Zoo
Bronx Zoo
Chattanooga Zoo
Dallas Zoo
Disney's Animal Kingdom
Fort Worth Zoo
Gladys Porter Zoo
Living Desert Zoo & Gardens
Omaha's Henry Doorly Zoo
Philadelphia Zoo
Riverbanks Zoo & Garden
San Antonio Zoo
San Diego Zoo Safari Park
Southwicks Zoo
Topeka Zoo
Tulsa Zoo
Zoo Atlanta
Zoo Tampa

Okapi (11 holders)
Bronx Zoo
Cincinnati Zoo & Botanical Garden
Dallas Zoo
Houston Zoo
Los Angeles Zoo & Botanical Garden
Omaha's Henry Doorly Zoo
San Antonio Zoo
San Diego Zoo
San Diego Zoo Safari Park
Sedgwick County Zoo
Zoo Tampa

Now obviously I am missing many, many holders for these animals. I likely will for most taxa I cover. But this is where you all come in, let's fill those gaps!

~Thylo
 
Hogle zoo had Giraffe when I went in July, I'm pretty sure they were just generic
 
Giraffes
Masai Giraffe (10 holders)
Brevard Zoo
Cincinnati Zoo & Botanical Garden
Franklin Park Zoo
Houston Zoo
Los Angeles Zoo & Botanical Gardens
Nashville Zoo
Roger Williams Park Zoo
San Diego Zoo
San Diego Zoo Safari Park
Santa Barbara Zoo

Rothschild's/Nubian Giraffe (suspected) (2 holders)
B. Bryan Preserve
San Diego Zoo Safari Park

Giraffe (sensu lato) (19 holders)
Audubon Species Survival Center
Audubon Zoo
Bronx Zoo
Chattanooga Zoo
Dallas Zoo
Disney's Animal Kingdom
Fort Worth Zoo
Gladys Porter Zoo
Living Desert Zoo & Gardens
Omaha's Henry Doorly Zoo
Philadelphia Zoo
Riverbanks Zoo & Garden
San Antonio Zoo
San Diego Zoo Safari Park
Southwicks Zoo
Topeka Zoo
Tulsa Zoo
Zoo Atlanta
Zoo Tampa

Okapi (11 holders)
Bronx Zoo
Cincinnati Zoo & Botanical Garden
Dallas Zoo
Houston Zoo
Los Angeles Zoo & Botanical Garden
Omaha's Henry Doorly Zoo
San Antonio Zoo
San Diego Zoo
San Diego Zoo Safari Park
Sedgwick County Zoo
Zoo Tampa

Now obviously I am missing many, many holders for these animals. I likely will for most taxa I cover. But this is where you all come in, let's fill those gaps!

~Thylo
Masai Giraffes are also kept by Columbus, Racine, Toledo, The Wilds, and (apparently) Timbavati Wildlife Park.

Saint Louis, Columbus, and Brookfield all keep Okapi.

Lincoln Park and The Wilds claims their Giraffes are pure Rothschild's, not sure if this is accurate.

Generic Giraffes are kept by Henry Vilas, NEW, Milwaukee, Como Park, Columbus, The Wilds, Saint Louis, Detroit, Binder Park, Indianapolis, Brookfield, and Timbavati Wildlife Park.
 
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Wild Cattle
American Bison (pure) (5 holders)
Bronx Zoo
Minnesota Zoo
Oakland Zoo
Oklahoma City Zoo
Sedgwick County Zoo

American Bison (impure) (6 holders)
Action Wildlife
Queens Zoo
Roger Williams Park Zoo
Rosamond Gifford Zoo
Space Farms Zoo & Museums
Wildlife World Zoo

Lowland Anoa (7 holders)
Fort Worth Zoo
Micanopy Zoological Preserve
Point Defiance Zoo & Aquarium
Rum Creek Ranch
San Diego Zoo Safari Park
Timbavati Wildlife Park
Zoo Tampa

Javan Banteng (2 holders)
Rum Creek Ranch
San Diego Zoo Safari Park

Gaur (6 holders)
Bronx Zoo
Gladys Porter Zoo
Lee G. Simmons Wildlife Safari Park
Living Treasures (not sure which one, there are two in Virginia but one has Gaur supposedly)
San Diego Zoo Safari Park
Zoo Miami

Cape Buffalo (2 holders)
Denver Zoo
San Diego Zoo Safari Park

Again, there are definitely others for these, especially the impure bison, it's just a little difficult to check every single zoo website at 11pm :p

~Thylo
 
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